Answer:
Option D, A red allele present on at least one of two homologous chromosomes
Explanation:
It is given that in a beetle species the red color is dominant over the white color. Thus, in order to have a offspring of red color at least one of the two homologous chromosome must have red allele.
This can be further understood from the cross below -
Suppose one of the parent is homozygous red with genotype "RR" and other parent is homozygous white with genotype "rr"
If a cross is carried out between the two parents, the following offsprings will result -
RR x rr
Rr, Rr, Rr, Rr
Since there is no co dominance, thus the dominant allele will express itself completely and hence at least one of the homologous chromosome of offspring must have red color allele.